Phase contrast image of biomass from the Tasmanian winery WWTP and corresponding composite FISH image: EUBmix (FLUOS) alone = green; EUBmix (FLUOS) + AMAR839 (CY3) = yellow. Scale bar = 10 microns. - Source:1Aerobic heterotroph
Tetrad forming organisms orignally believed to be GAOs. However pure cultures were shown to be unable to assimilate carbon under anaerobic conditions 2. Aerobic heterotrophs, which have a high capacity for PHA storage 2. No information on their in situ physiology in activated sludge is availabe.
Has been associated with non-filamentous viscous bulking in industrial plants 3.
 McIlroy SJ (2010) Phylogenetic Diversity and Ecophysiology of Alphaproteobacterial Glycogen Accumulating Organisms in Enhanced Biological Phosphorus Removal Activated Sludge Systems. PhD Thesis La Trobe University. - Mcilroy -
 Falvo, Levantesi, Rossetti, Seviour, Tandoi - Synthesis of intracellular storage polymers by Amaricoccus kaplicensis, a tetrad forming bacterium present in activated sludge. - J. Appl. Microbiol. 91(2): 299-305.
 McIlroy, Speirs, Tucci, Seviour - In situ profiling of microbial communities in full-scale aerobic sequencing batch reactors treating winery waste in australia. - Environ. Sci. Technol. 45(20): 8794-803. doi:10.1021/es2018576
 Maszenan, Seviour, Patel, Wanner - A fluorescently-labelled r-RNA targeted oligonucleotide probe for the in situ detection of G-bacteria of the genus Amaricoccus in activated sludge. - J. Appl. Microbiol. 88(5): 826-35.
 Maszenan, Seviour, Patel, Rees, McDougall - Amaricoccus gen. nov., a gram-negative coccus occurring in regular packages or tetrads, isolated from activated sludge biomass, and descriptions of Amaricoccus veronensis sp. nov., Amaricoccus tamworthensis sp. nov., Amaricoccus macauensis sp. nov., and Amaricoccus kaplicensis sp. nov. - Int. J. Syst. Bacteriol. 47(3): 727-34. doi:10.1099/00207713-47-3-727
 Blackall, Rossetti, Christensson, Cunningham, Hartman, Hugenholtz, et al. - The characterization and description of representatives of 'G' bacteria from activated sludge plants. - Lett. Appl. Microbiol. 25(1): 63-9.